SUBGENUS Pleurothallis SECTION Macrophyllae-Fasciculatae Lindl. 1859

Characterized by the single flowered inflorescence with the peduncle hidden within a spathe, the usually short pedicels, the entire, fringed to pubescent petals that are not sigmoid in outline and a thick, obtuse, oblong to transverse lip and a shorter, broader column with an apical anther and stigma that is almost always bilobed. This subsection and amphygiae are not distinctly separated in various borderline species and for this reason they later were separated by Luer into the genus Acronia [2005] with three sections, Acronia, Amphygia and Macrophyllae-fasciculatae. TYPE Species for the SECTION

!Pleurothallis calceolaris Rchb.f. 1876

1. Pleurothallis acestrophylla Luer 1975 Ecuador and Peru

2. Pleurothallis acutilabia Luer 1999 Colombia

3. Pleurothallis adeleae Luer 1981 Ecuador and Ecuador

4. Pleurothallis adelphe Luer & Hirtz 1996 Ecuador

5. Pleurothallis adonis Luer 1976 Colombia and Ecuador

6. Pleurothallis adventurae Karremans & Bogarín 2011 Costa Rica

7. Pleurothallis aguirrei Luer & R.Escobar 2003 Colombia

8. Pleurothallis alopex Luer 1976 Bolivia and Paraguay

9. Pleurothallis altimonile Luer & R.Escobar 1981 Colombia

10. Pleurothallis ambyx Luer & R.Escobar 1981 Colombia and Ecuador

11. Pleurothallis andreaskayi Mark Wilson & B.T. Larsen 2022 Colombia, Ecuador

12. Pleurothallis andreettae (Luer) J.M.H.Shaw 2014 Ecuador

13. Pleurothallis angusta Ames & C.Schweinf. 1925 Costa Rica and Panama

14. Pleurothallis angustissima (Luer & Hirtz) J.M.H.Shaw 2014 Ecuador

15. Pleurothallis anthurioides A.Doucette 2016 Panama

16. Pleurothallis apopsis Luer 1979 Ecuador

17. Pleurothallis applanata Luer & Dalström 1996 Ecuador

18. Pleurothallis arctata Luer 1997 Peru

19. Pleurothallis ascera Luer & R. Escobar 1996 Costa Rica, Panama and Colombia

20. Pleurothallis asplundii Luer 1996 Ecuador

21. Pleurothallis aurita Schweinf 1940 Costa Rica

22. Pleurothallis baezensis (Luer & Hirtz) J.M.H.Shaw 2014 Ecuador

23. Pleurothallis barrowii Schuit. 2018 Ecuador?

24. Pleurothallis batrachus Luer & Hirtz 1996 Ecuador

25. Pleurothallis baudoensis Luer & R.Escobar 1998 Colombia

26. Pleurothallis belocardia Schltr. 1924 Colombia

27. Pleurothallis bilobulata M.M.Jiménez, Ocupa & Vélez-Abarca 2021 Ecuador

28. Pleurothallis bitumida Luer 1996 Costa Rica

29. Pleurothallis bivalvis Lindl. 1846 Ecuador, Colombia and Venezuela

30. Pleurothallis blepharopetala Schltr.1915 Ecuador

31. Pleurothallis bogarinii Pupulin & J.D. Zúñiga 2007 Costa Rica

32. Pleurothallis bothros Luer 1996 Costa Rica

33. Pleurothallis bovilingua Luer & R.Escobar 1998 Colombia

34. Pleurothallis braidiana [Luer] in ed. without location

35. Pleurothallis bucculenta Luer 1976 Ecuador

36. Pleurothallis bulbosa [Luer & Escobar] Pfahl 2012 Colombia

37. Pleurothallis calamifolia Luer & R.Escobar 1996 Colombia

38. !Pleurothallis calceolaris Rchb.f. 1876 Colombia

39. Pleurothallis callifera C.Schweinf. 1951 Venezuela

40. Pleurothallis calogramma Luer 2003 Colombia

41. Pleurothallis callosa Diaz & Pupulin 2021 Costa Rica

42. Pleurothallis calolalax Luer & R.Escobar 1998 Colombia

43. Pleurothallis canaligera Rchb.f 1877 Colombia, Ecuador and Peru

44. Pleurothallis canidentis Luer & R. Escobar 1998 Costa Rica, Panama and Colombia

45. Pleurothallis cardiochila L.O. Williams 1942 Costa Rica and Panama

46. Pleurothallis cardiocrepis Rchb.f.1888 Peru?

47. Pleurothallis cardiophylax Rchb.f. 1888 Colombia, Ecuador and Peru

48. Pleurothallis cardiostola Rchb. f. 1854 Colombia, Venezuela, Peru, Bolivia and Ecuador

49. Pleurothallis cardiothallis Rchb.f 1857 Mexico, Belize, Costa Rica, Nicaragua, Colombia and Ecuador

50. Pleurothallis carduela (Luer) J.M.H.Shaw 2014 Ecuador

51. Pleurothallis carmensotoana Mark Wilson & B.T. Larsen 2022 Ecuador, Peru

52. Pleurothallis carnosa Braid 1924 Colombia

53. Pleurothallis carrenoi Carnevali & I.Ramírez 1987 Venezuela

54. Pleurothallis cassidata Luer & R.Escobar 1998 Colombia

55. Pleurothallis castanea Mark Wilson, G.Merino & J.D.Werner 2016 Ecuador

56. Pleurothallis caucensis Mark Wilson 2017 Colombia

57. Pleurothallis cauda-hirundinis Luer & J.Portilla 2002 Ecuador

58. Pleurothallis cedrinorum Luer & Dalström 1996 Ecuador

59. Pleurothallis chama Luer 1977 Ecuador

60. Pleurothallis chaparensis (Luer & R.Vásquez) J.M.H.Shaw 2014 Bolivia

61. Pleurothallis chavezii Luer 1996 Costa Rica

62. Pleurothallis chicalensis M.M.Jiménez & Baquero 2018 Ecuador

63. Pleurothallis cobriformis L.O.Williams 1940 Panama

64. Pleurothallis complanata Luer & Hirtz 1996 Ecuador

65. Pleurothallis compressa Luer 1996 Costa Rica

66. Pleurothallis conformalis Luer & Dalström 2003 Ecuador

67. Pleurothallis conicostigma Luer 1975 Ecuador

68. Pleurothallis constricta Luer & R.Escobar R. 1998 Colombia

69. Pleurothallis convexa Luer & Hirtz 1996 Ecuador

70. Pleurothallis cordata (Ruiz & Pav.) Lindl. 1830 Colombia, Venezuela, Ecuador, Peru and Bolivia

71. Pleurothallis cordifolia Rchb.f. & Wagener 1854 Costa Rica, Panama, Venzuela, Colombia and Ecuador

72. Pleurothallis coriacardia Rchb. f. 1854 Ecuador and Peru

73. Pleurothallis correllii Luer 1976 Guatemala, El Salvador and Mexico

74. Pleurothallis corysta Luer 1978 Ecuador

75. Pleurothallis crateriformis C.Schweinf. 1951 Ecuador and Peru

76. Pleurothallis crossota Luer & Dalström 1996 Ecuador

77. Pleurothallis crucifera Luer & Hirtz 1996 Ecuador

78. Pleurothallis culpameae (Luer) J.M.H.Shaw 2014 Ecuador

79. Pleurothallis cuzcoensis [Luer] ined. Peru

80. Pleurothallis cutucuensis Luer 1977 Ecuador

81. Pleurothallis cyanea Luer & R. Escobar 1984 Colombia

82. Pleurothallis dariensis (Kolan. & Szlach.) Bogarín 2014 Panama

83. Pleurothallis deflexa Luer 1975 Ecuador

84. Pleurothallis dejavu Luer & Hirtz 1988Ecuador

85. Pleurothallis depressa Luer & R. Escobar 1996 Colombia

86. Pleurothallis dewildei Luer & R.Escobar 1998 Colombia

87. Pleurothallis diabolica Luer & R. Escobar 1981 Colombia Ecuador

88. Pleurothallis diazii (Luer & Endara) J.M.H.Shaw 2014 Peru

89. Pleurothallis dibolia Luer 1975 Ecuador

90. Pleurothallis dilemma Luer 2001 Ecuador

91. Pleurothallis discoidea Lindl. 1835 Trinidad, Venezuela, Surinam, French Guiana, Ecuador and Peru

92. Pleurothallis dorotheae Luer 1980 Costa Rica

93. Pleurothallis dracuncula (Luer & Hirtz) J.M.H.Shaw 2014 Ecuador

94. Pleurothallis drewii Luer 2000

    Ecuador

95. Pleurothallis dubbeldamii Luer 2003 No Collection Data

96. Pleurothallis ensata Luer 1975 Ecuador

97. Pleurothallis epiglottis Luerb 1980 Ecuador

98. Pleurothallis erythrium Luer 1978 Ecuador

99. Pleurothallis escobarii (Luer) J.M.H.Shaw 2014 Colombia

100. Pleurothallis excavata Schltr. 1923 Costa Rica and Panama

101. Pleurothallis fantastica Ames 1936 Costa Rica

102. Pleurothallis folsomii (Luer & Endara) J.M.H.Shaw 2014 Panama

103. Pleurothallis fonnegrae (Luer & Endara) J.M.H.Shaw 2014 Colombia

104. Pleurothallis fossulata Luer & R.Escobar 1998 Colombia and Ecuador

105. Pleurothallis franciana Sierra-Aziz 2023 Colombia

106. Pleurothallis galerita Luer 1981 Ecuador

107. Pleurothallis ganymedes Luer & R.Escobar 1981 Colombia

108. Pleurothallis gargantua Luer 1996 Ecuador

109. Pleurothallis geographica Luer 1977 Ecuador

110. Pleurothallis gigiportillae A.Doucette & J.Portilla 2016 Ecuador

111. Pleurothallis giraffa Luer 1979 Ecuador

112. Pleurothallis giraldoi Luer 1999 Colombia

113. Pleurothallis glabra Luer & R.Escobar 1996 Colombia

114. Pleurothallis globosa Luer & R.Escobar 1996 Colombia

115. Pleurothallis gonzaleziorum Pupulin, Diaz & Pridgeon 2021 Costa Rica

116. Pleurothallis gonzaloi J.S. Moreno, Rinc.-González & Gal.-Tar 2022 Colombia

117. Pleurothallis grandiflora Lindl. 1835 Venezuela, Colombia, Ecuador, Peru and Bolivia

118. Pleurothallis grandilingua Pupuplin, M Diaz and Pridgeon 2021 Costa Rica

119. Pleurothallis grobleri (Luer & Hirtz) J.M.H.Shaw 2014 Ecuador

120. Pleurothallis hemileuca Luer 1982 Panama

121. Pleurothallis hoeijeri Luer & Hirtz 1988 Ecuador

122. Pleurothallis homalantha Schltr.1923 El Salvador, Honduras, Nicaragua, Costa Rica, Panama and Brazil

123. Pleurothallis imperialis Luer 1975 Ecuador and Colombia

124. Pleurothallis index Luer 1980 Ecuador and Colombia

125. Pleurothallis isthmica Luer 1977 Honduras, Costa Rica and Panama

126. Pleurothallis jaramilloi Luer 2001 Colombia

127. Pleurothallis jupiter Luer 1975 Ecuador

128. Pleurothallis kashi-menkakarai Mashendo-Jimbicti, Vélez-Abarca & M.M.Jiménez 2023 Ecuador

129. Pleurothallis killipii Garay 1956 Colombia

130. Pleurothallis lacera Luer 1975 Colombia and Ecuador

131. Pleurothallis lanigera Luer & Hirtz 1988 Ecuador

132. Pleurothallis lapoi M.M.Jiménez & Vélez-Abarca 2023 Ecuador

133. Pleurothallis latipetala Garay 1952 Colombia

134. Pleurothallis lemniscifolia Luer 1975 Ecuador

135. Pleurothallis leucantha Schltr. 1912 Guatemala

136. Pleurothallis lilijae Foldats 1968 Venezuela, Colombia and Ecuador

137. Pleurothallis linguifera Lindl. 1859 Colombia, Ecuador, Peru, Bolivia and Venezuela

138. Pleurothallis llanganatensis (Luer & Hirtz) J.M.H.Shaw 2014

139. Pleurothallis longipetala Bogarin & Belfort 2021 Costa Rica

140. Pleurothallis loreae Carnevali & I.Ramírez 2003 Venezuela

141. Pleurothallis luna-crescens Pupulin, J.Aguilar & Mel.Fernández 2017 Costa Rica

142. Pleurothallis lunaris Luer & R.Escobar 1981 Colombia

143. Pleurothallis lynniana [Luer] Comb. novo. Pfahl 2012 Ecuador

144. Pleurothallis macra Lindl. 1859 Ecuador and Peru

145. Pleurothallis macrocardia Rchb.f. 1855 Colombia, Ecuador and Bolivia

146. Pleurothallis maduroi Luer 1997 Panama

147. Pleurothallis magnipetala C.Schweinf. 1951 Peru

148. Pleurothallis mahechae J.S. Moreno, Sierra-Ariza & L.C. Pina 2022 Colombia

149. Pleurothallis marioandresavilae Vélez-Abarca & M.M.Jiménez 2023 Ecuador, Peru

150. Pleurothallis marioi Zambrano & Solano 2017 Ecuador

151. Pleurothallis marthae Luer & R. Escobar 1996 Colombia

152. Pleurothallis mastodon Luer & Hirtz 1996 Ecuador

153. Pleurothallis matudana C. Schweinf. 1938 Mexico, Guatemala, Honduras, Nicaragua, Ecuador and Bolivia

154. Pleurothallis megaglossa Luer & Dalström 1996 Ecuador

155. Pleurothallis megalorhina Luer & R.Escobar 1996 Colombia

156. Pleurothallis mesopotamica Bogarin & Belfort 2021 Costa Rica

157. Pleurothallis microcardia Rchb.f. 1855 Colombia, Venezuela, Ecuador, Peru and Bolivia

158. Pleurothallis millei Schltr. 1915 Ecuador

159. Pleurothallis miniatura (Luer, Thoerle & F.Werner) J.M.H.Shaw 2014 Ecuador

160. Pleurothallis minutilabia Mark Wilson, Tobar & Á.J.Pérez 2018 Ecuador

161. Pleurothallis muriculata Luer & Hirtz 1996 Ecuador

162. Pleurothallis navisepala Pupulin, J.Aguilar & M.Díaz 2017 Costa Rica

163. Pleurothallis neobarbosae J.M.H.Shaw 2014 Peru

164. Pleurothallis neorinkei A.Doucette 2016 Colombia

165. Pleurothallis nephroglossa Schltr. 1921 Ecuador

166. Pleurothallis nipterophylla C.A.Luer 1976 Ecuador

167. Pleurothallis nitida Luer 1977 Panama

168. Pleurothallis nossax Luer & R. Escobar 1981 Colombia

169. Pleurothallis octavioi Luer & R.Escobar R. 1981 Colombia

170. Pleurothallis omoglossa Luer 1976 Ecuador

171. Pleurothallis oncoglossa Luer 1996 Costa Rica

172. Pleurothallis ortegae Luer & Hirtz 1996 Ecuador

173. Pleurothallis orygmoglossa (Luer & Dressler) J.M.H.Shaw 2014 Panama

174. Pleurothallis oscarii Archila & Chiron 2011 Guatemala

175. Pleurothallis oxapampae Luer 1982 Peru

176. Pleurothallis palliolata Ames 1922 Costa Rica and Panama

177. Pleurothallis pansamalae Schltr. 1912 Mexico, Guatemalla, Belize, El Salvador, Honduras and Nicaragua

178. Pleurothallis papillingua A.Doucette & J.Portilla 2016 Ecuador

179. Pleurothallis paquishae Luer 1996 Ecuador and Peru

180. Pleurothallis peculiaris Luer 1976 Panama

181. Pleurothallis pelicophora Luer 2001 Ecuador

182. Pleurothallis penelops Luer 1978 Ecuador

183. Pleurothallis perijaensis Dunst. 1978 Venezuela, Colombia and Ecuador

184. Pleurothallis peroniocephala Luer 1975 Ecuador

185. Pleurothallis perryi Luer 1979 Ecuador

186. Pleurothallis persimilis Luer & Hirtz 2003 Ecuador

187. Pleurothallis petroana Sierra-Ariza 2023 Colombia

188. Pleurothallis phyllocardia Rchb.f. 1866 Costa Rica and Panama

189. Pleurothallis phyllocardioides Schltr. 1923 Guatemala, Nicaragua, Costa Rica, Panama, Colombia, Ecuador, Peru, Bolivia and Venezuela

190. Pleurothallis phymatodea Luer 1976 Ecuador

191. Pleurothallis pileata Luer & R.Escobar 1982 Colombia

192. Pleurothallis platypetala Luer & R.Escobar 1996 Colombia

193. Pleurothallis platysepala Schltr. 1920 Colombia

194. Pleurothallis portillae Luer 1979 Colombia and Ecuador

195. Pleurothallis prolaticollaris Luer 1977 Ecuador

196. Pleurothallis pudica Pupulin, J.Aguilar & M.Díaz 2017 Costa Rica

197. Pleurothallis pyelophera [Luer] comb. novo. Pfahl 2013 Ecuador

198. Pleurothallis quitu-cara Carrera & Baquero 2018 Ecuador

199. Pleurothallis radula Luer 1979 Costa Rica

200. Pleurothallis ramiromedinae Thoerle & Hirtz 2018 Colombia, Ecuador

201. Pleurothallis rectipetala Ames & Schweinf. 1925 Panama

202. Pleurothallis recurvata Luer & Hirtz 1996 Ecuador

203. Pleurothallis reginae Garay 1953 Colombia

204. Pleurothallis reptans Luer 1981 Ecuador

205. Pleurothallis rhinocera (Luer & Sijm) J.M.H.Shaw 2013 Ecuador

206. Pleurothallis rhodoglossa Schltr.1922 Costa Rica, Panama, Colombia and Ecuador

207. Pleurothallis rikseniana Mark Wilson & B.T. Larsen 2022 ?Ecuador?

208. Pleurothallis ripleyi Luer 1979 Ecuador

209. Pleurothallis ruberrima Lindley 1846 Colombia, Ecuador, Peru, Bolivia and Venezuela

210. Pleurothallis rubrifolia Mark Wilson, Tobar & Salas Guerr. 2016 Ecuador, Peru

211. Pleurothallis rubroinversa Luer 1975 Ecuador

212. Pleurothallis rugosa Luer & R.Escobar 1982 Colombia

213. Pleurothallis sabanillae M.M.Jiménez & Vélez-Abarca 2023 Ecuador

214. Pleurothallis sagittilabia Luer 1998 Colombia

215. Pleurothallis sanchoi Ames 1923 Guatemala, Costa Rica and Panama

216. Pleurothallis sannio Luer & R.Escobar 1988 Colombia

217. Pleurothallis sarcochila Garay 1953 Ecuador

218. Pleurothallis scabrilinguis Lindl. 1836 Colombia Ecuador and Peru

219. Pleurothallis scaphipetala Luer 1996 Costa Rica

220. Pleurothallis scotinantha Pupulin, M.Díaz & J.Aguilar 2017 Costa Rica

221. Pleurothallis scurrula Luer 1976 Peru

222. Pleurothallis semiscabra Lindl. 1846 Venezuela

223. Pleurothallis sergioi Luer & R.Escobar 2003 Colombia

224. Pleurothallis serricardia Schltr. 1924 Colombia

225. Pleurothallis sigynes Luer 1976 Colombia & Ecuador

226. Pleurothallis silvae-pacis Karremans 2011 Costa Rica

227. Pleurothallis siphoglossa Luer & R. Escobar 1996 Colombia

228. Pleurothallis sotarae Schltr. 1920 Colombia

229. Pleurothallis sphaerantha Luer 1975 Ecuador

230. Pleurothallis stelidilabia Luer 1979 Ecuador

231. Pleurothallis stellata I.Ramírez & Carnevali 2003 Venezuela

232. Pleurothallis stenosepala Rolfe 1892 Venezuela, Colombia, Ecuador, Peru and Bolivia

233. Pleurothallis stenota Luer 1977 Colombia and Ecuador

234. Pleurothallis subtilis C.Schweinf. 1951 Venezuela and Colombia

235. Pleurothallis x subversa Pupulin & Bogarin 2021 Costa Rica

236. Pleurothallis tamaensis Foldats 1968 northwestern Venezuela

237. Pleurothallis tapatiensis Pupulin, M Diaz & Pridgeon 2021 Costa Rica

238. Pleurothallis teaguei Luer 1996 Ecuador

239. Pleurothallis tinajillensis M.M.Jiménez, H. Garzón & Vélez-Abarca 2023 Ecuador

240. Pleurothallis titan Luer 1977 Panama and Colombia

241. Pleurothallis tobarii [Luer & Hirtz] comb. novo. Pfahl 2012 Ecuador

242. Pleurothallis tonduzii Schltr. 1918 Costa Rica, Panama and Peru?

243. Pleurothallis trachysepala Kraenzl. 1905 Colombia, Ecuador, Peru

244. Pleurothallis tragulosa Luer & Hirtz 1996 Ecuador

245. Pleurothallis transversilabia Carnevali & I.Ramírez 2000 Venezuela

246. Pleurothallis triangulilabia C.Schweinf. 1937 Costa Rica

247. Pleurothallis tridentata Klotzsch 1840 Venezuela, Colombia, Ecuador, Peru? and Bolivia?

248. Pleurothallis troglodytes Luer 1982 Ecuador

249. Pleurothallis tyria Luer & Hirtz 1996 Ecuador

250. Pleurothallis undulata Poepp. & Endl. 1836 Peru

251. Pleurothallis upanoensis Luer & Hirtz 2003 Ecuador

252. Pleurothallis valladolidensis Luer 1983 Ecuador

253. Pleurothallis variabilis Luer 1979 Colombia and Ecuador

254. Pleurothallis villahermosae Sierra-Ariza, Rinc.-González & Villanueva 2022 Colombia

255. Pleurothallis vinealis Luer & R.Escobar 1986 Colombia and Ecuador

256. Pleurothallis volans Luer & Hirtz 1996 Ecuador

257. Pleurothallis whitteniana MWilson & B T Larsen 2022 Ecuador, Peru

258. Pleurothallis zarumae Luer & Hirtz 1996 Ecuador

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